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Article title
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Abstract
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Key words
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Introduction
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Materials and methods
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Sampling
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Phylogenetic analyses
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Molecular dating
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Morphological measurements and analyses
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Results
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Phylogenetic analyses
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Morphological analyses
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Taxonomic account
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Discussion
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Acknowledgements
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Additional information
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Conflict of interest
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Ethical statement
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Funding
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Author contributions
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Author ORCIDs
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Data availability
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References
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Appendix 1
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Supplementary materials
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